UC Davis Department of Entomology

Research in the Carey Laboratory

     Research in my laboratory focuses on biodemography: an emerging division of classical demography that brings life table techniques, mortality models, experimental systems, and comparative methods to bear on questions concerned with the fundamental determinants of mortality, longevity, aging and life span. I am interested in insect mortality dynamics, mortality trajectories, and the evolution of life span and extended longevity. I am also interested in aging with special emphasis on the oldest individuals in a population, and with studying the behavioral and social components of life span and aging. The evolution of social organization as a component of life history is an important research interest in my laboratory. While many of our studies are on captive populations, we are moving into the study of aging in nature; attempting to determine mortality dynamics in field conditions.

Ph.D. Students and Postdoctoral Scientists in the Laboratory

Address | Interests | Natural History Data | Education | Publications

Freerk Molleman
[Postdoctorate Research Associate]
Office: 69 Briggs Hall
Phone: (530) 754-4872
Email: fmolleman@ucdavis.edu
freerkmolleman@hotmail.com
Fax: (530) 752-1537

Advisor: James R. Carey

Department of Entomology
UC Davis
One Shields Avenue
Davis, CA 95616-8584
USA

Field-site:
Makerere University Biological Field Station
P.O.Box 409
Fort Portal
Uganda

Interests
The Introduction to my Ph. D. thesis gives an impression of my interests:
Biodiversity, life-history, evolution, ecology

I am happy to work as a post doc with James Carey within the Biodemographic determinants of life span program.
Biodemography combines demography with the biology of aging and an evolutionary approach. My research focuses on life-span evolution in fruit-feeding butterflies in Uganda and can be seen as a continuation of my Ph. D. work in Leiden. Life-span is a central life history trait and life history theory is central to biology, so I am right in the middle of the jungle, ready to get lost. I am also involved in a project on the demography of impairment.

Click Here for my PDF on the "Aspects of biodiversity connected by arrows of interest to evolutionary biology".

(Bonny, Easter and Moses at Lake Victoria)

'Bonny Balyeganira and Moses Musana work for Kibale butterfly project in Uganda. I taught them to identify fruit-feeding butterflies in the field and they can independently perform various experiments and surveys on butterflies. Easter has been working on data entry. Sometimes we go with the team on a trip.'

Conservation and Development

Charaxes kahldeni feeding on civet dung)

Conservation and development are two much related subjects that have my interest. Some of my research has applied sides, such as the biodiversity monitoring, and the project I did with my wife, Gosia on human-wildlife conflict in a rainforest in Cameroon. In Uganda we are active in education and nature education. The foundation that Gosia and I started is providing schools and local libraries with books. Have a look at the web-site! http://www.stichtingkeo.nl/

Natural History Data

(Euphaedra eusemoides laying eggs on Uvariopsis congolensis)

I use this web-site for publishing natural history data. Under Construction

Nearly complete species list of fruit-feeding Nymphalids from Kibale Forest, Uganda and some other butterflies, illustrated with pictures from the wild and in some cases mounted individuals or caterpillars. Click here

(Catuna crithea in copula)

Click here for list of butterfly species that I found at the northern periphery of the Dja Faunal Reserve in Cameroon. A complete list with photographs of mounted specimens is Under Construction.

Education:

16 December 2004 Successful defense of Ph.D. thesis

April 1999-July 2003 Ph.D. study Department of Evolutionary Biology Leiden University, The Netherlands Graduate School Functional Ecology

Advisors Professor Paul M. Brakefield and Dr. Bas J. Zwaan

Title Patterns of Biodiversity and Life History evolution in fruit-feeding butterflies

Sept 1993-Sept 1998 Masters study Biology, Orientation Population/Ecology
Wageningen Agricultural University

Subjects Evolutionary ecology, Advanced statistics, Computational statistics, Population Dynamics

Publications:

Molleman, F., Krenn, H., Brakefield, P. M., DeVries, P. J. Zwaan, B. J. Food intake of fruit-feeding butterflies: evidence for adaptive variation in proboscis morphology (In Press Biological Journal of the Linnean Society)

Molleman, F., Grunsven, R. van, Liefting, M., Brakefield, P. M., Zwaan, B. J. Is male puddling behaviour of tropical butterflies targeted at sodium for nuptial gifts or for activity? (In Press Biological Journal of the Linnean Society)

Molleman, F, Kop, A., Brakefield, P. M., DeVries, P. J., Zwaan, B. J. Temporal and vertical patterns in biodiversity of fruit-feeding butterflies in Uganda (In Press Biodiversity and Conservation)

Molleman, F., Alphen, M. van, Brakefield, P. M., Zwaan, B. J. Food quality and preferences for fruit-feeding butterflies in Kibale Forest, Uganda (In Press Biotropica)

Molleman, F., Hecq J. Host-plant records and photographs of immature Euphaedra from Kibale National Park, Uganda (Lepidoptera Nymphalidae) (In Press Lambillionea)

Molleman, F., Zwaan, B. J., Brakefield, P. M. 2004 The effect of male sodium diet and mating history on female reproduction in the puddling butterfly Bicyclus anynana. Behavioural Ecology and Sociobiology. 56: 404-411

Molleman, F., Walter, D. E. 2001 Niche segregation and can-openers: Scydmaenid beetles as predators of armoured mites in Australia. Acaralogy: Proceedings of the 10th International Congress pp 283-288

Molleman, F., Drukker, B., Blommers, L., 1997 A trap for monitoring pear psylla predators using dispensers with the synomone methylsalicylate. Proceedings of the section Experimental and Applied Entomology of the Netherlands Entomological Society (N.E.V.) 8: pp177-182

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Department of Entomology, UC Davis, One Shields Avenue, Davis, CA 95616-8584     phone: (530) 752-0475     fax: (530) 752-1537

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This page last updated:    April 03, 2007

Freerk Molleman [Postdoctorate Research Associate] Office: 69 Briggs Hall Phone: (530) 754-4872 Email: fmolleman@ucdavis.edu freerkmolleman@hotmail.com Fax: (530) 752-1537
Advisor: James R. Carey Department of Entomology UC Davis One Shields Avenue Davis, CA 95616-8584 USA Field-site: Makerere University Biological Field Station P.O.Box 409 Fort Portal Uganda Interests The Introduction to my Ph. D. thesis gives an impression of my interests: Biodiversity, life-history, evolution, ecology I am happy to work as a post doc with James Carey within the Biodemographic determinants of life span program. Biodemography combines demography with the biology of aging and an evolutionary approach. My research focuses on life-span evolution in fruit-feeding butterflies in Uganda and can be seen as a continuation of my Ph. D. work in Leiden. Life-span is a central life history trait and life history theory is central to biology, so I am right in the middle of the jungle, ready to get lost. I am also involved in a project on the demography of impairment. Click Here for my PDF on the "Aspects of biodiversity connected by arrows of interest to evolutionary biology".
	(Bonny, Easter and Moses at Lake Victoria)
'Bonny Balyeganira and Moses Musana work for Kibale butterfly project in Uganda. I taught them to identify fruit-feeding butterflies in the field and they can independently perform various experiments and surveys on butterflies. Easter has been working on data entry. Sometimes we go with the team on a trip.'
Conservation and Development
	Charaxes kahldeni feeding on civet dung)
Conservation and development are two much related subjects that have my interest. Some of my research has applied sides, such as the biodiversity monitoring, and the project I did with my wife, Gosia on human-wildlife conflict in a rainforest in Cameroon. In Uganda we are active in education and nature education. The foundation that Gosia and I started is providing schools and local libraries with books. Have a look at the web-site! http://www.stichtingkeo.nl/
Natural History Data
	(Euphaedra eusemoides laying eggs on Uvariopsis congolensis)
I use this web-site for publishing natural history data. Under Construction Nearly complete species list of fruit-feeding Nymphalids from Kibale Forest, Uganda and some other butterflies, illustrated with pictures from the wild and in some cases mounted individuals or caterpillars. Click here
	(Catuna crithea in copula)
Click here for list of butterfly species that I found at the northern periphery of the Dja Faunal Reserve in Cameroon. A complete list with photographs of mounted specimens is Under Construction.

Education:
16 December 2004	Successful defense of Ph.D. thesis
April 1999-July 2003	Ph.D. study Department of Evolutionary Biology Leiden University, The Netherlands Graduate School Functional Ecology
Advisors	Professor Paul M. Brakefield and Dr. Bas J. Zwaan
Title	Patterns of Biodiversity and Life History evolution in fruit-feeding butterflies
Sept 1993-Sept 1998	Masters study Biology, Orientation Population/Ecology Wageningen Agricultural University
Subjects	Evolutionary ecology, Advanced statistics, Computational statistics, Population Dynamics
Publications: Molleman, F., Krenn, H., Brakefield, P. M., DeVries, P. J. Zwaan, B. J. Food intake of fruit-feeding butterflies: evidence for adaptive variation in proboscis morphology (In Press Biological Journal of the Linnean Society) Molleman, F., Grunsven, R. van, Liefting, M., Brakefield, P. M., Zwaan, B. J. Is male puddling behaviour of tropical butterflies targeted at sodium for nuptial gifts or for activity? (In Press Biological Journal of the Linnean Society) Molleman, F, Kop, A., Brakefield, P. M., DeVries, P. J., Zwaan, B. J. Temporal and vertical patterns in biodiversity of fruit-feeding butterflies in Uganda (In Press Biodiversity and Conservation) Molleman, F., Alphen, M. van, Brakefield, P. M., Zwaan, B. J. Food quality and preferences for fruit-feeding butterflies in Kibale Forest, Uganda (In Press Biotropica) Molleman, F., Hecq J. Host-plant records and photographs of immature Euphaedra from Kibale National Park, Uganda (Lepidoptera Nymphalidae) (In Press Lambillionea) Molleman, F., Zwaan, B. J., Brakefield, P. M. 2004 The effect of male sodium diet and mating history on female reproduction in the puddling butterfly Bicyclus anynana. Behavioural Ecology and Sociobiology. 56: 404-411 Molleman, F., Walter, D. E. 2001 Niche segregation and can-openers: Scydmaenid beetles as predators of armoured mites in Australia. Acaralogy: Proceedings of the 10th International Congress pp 283-288 Molleman, F., Drukker, B., Blommers, L., 1997 A trap for monitoring pear psylla predators using dispensers with the synomone methylsalicylate. Proceedings of the section Experimental and Applied Entomology of the Netherlands Entomological Society (N.E.V.) 8: pp177-182

Administration \| Department \| Faculty \| Undergrad \| Graduate \| Alumni \| Course Descriptions \| Seminars \| FAQ's \| Search \| Web Links \| Home
Department of Entomology, UC Davis, One Shields Avenue, Davis, CA 95616-8584 phone: (530) 752-0475 fax: (530) 752-1537